(1Institute of Biotechnology and Germplasm Resources,Yunnan Academy of Agricultural Sciences,Key Lab of Agricultural Biotechnology of Yunnan Province,Key Lab of Southwestern Crop Gene Resources and Germplasm Innovation,Ministry of Agriculture,Kunming 650233,Yunnan,China;2Life Science College,Southwest Forestry University,Kunming 650224,Yunnan,China;3Tranditional Chinese Medicine College,Yunnan University of Tranditional Chinese Medicine,Kunming 650500,Yunnan,China)